as well as of the larva and pupa. There is a membranous lobe or jugum near the base of the wing, and the neuration of the hindwing is closely like that of the forewing, the radial nervure being five branched in both. The pupa has four or five movable segments, and 1 the larval prolegs have complete circles of hooklets. The three families of the Iugatae are not very closely related to each other. The Micropterygidae (often known as Eriocephalidae), comprising a few small moths with metallic wings, are the most primitive of all Lepidoptera. They are provided with functional mandibles, while the maxillae have distinct laciniae, well-developed palps and galeae not modified for suction (see fig. 3). The larva is remarkable on account of its long feelers, the presence of pairs of jointed prolegs on the first eight abdominal segments, an anal sucker beneath the last segment and bladder-like outgrowths on the cuticle. These curious larvae feed on wet moss. The family has only a few genera scattered widely over the earth's surface (Europe, America, Australia, New Zealand).

The Eriorraniidae resemble the Micropterygidae in appearance, but the imago has no mandibles, and the maxillae, though short and provided with conspicuous pal s, have no laciniae and form a proboscis as in Lepidoptera generality. The abdomen of the female carries a serrate piercing process, and the eggs are laid in the leaves of deciduous trees, the white larvae, with aborted legs, mining in the leaf tissue. The fully-fed larva winters in an underground cocoon and then changes into the most remarkable of all known lepidopterous pupae, with relatively enormous toothed mandibles which bite a way out of the cocoon in preparation for the final change. These pupal mandibles of the Eriocraniidae, together with the nature of the rmaginal maxillae in the Micropterygidae (Eriocephalidae) and the wing-neuration in both families, point strongly to a relationship between the Lepidoptera and the Trichoptera. The Hepialidae or swift moths-the third famil of the '[ugataeare in some respects specialized. The moths are ofy large or moderate size with the maxillae in a vestigial condition, no food being taken after the attainment of the perfect state. The larvae (fig. 12) feed either on roots or in' the wood of trees and shrubs, not attaining their growth in less than a year and some large exotic species living for two or three. The family is world-wide in range, and Australia possesses some almost gigantic and strangely coloured genera. Tineides.

A large assemblage of moths, mostly of small size, are included in this group. The wings have no jugum, but there is a frenuluni on the hindwing, which has, as in all the groups above the jugatae, only a single radial nervure. Three anal nervures are present in the hindwing in those families whose wings are well developed, but in several families of small moths the wings of both pairs are ver narrow and pointed, and the neuration is consequently reduced? The sub-costal nervure of the hindwing is usually present and distinct from the radial nervure. The egg is fiat except in the Cossidae and Castniidae in which it is upright. The larval prolegs, with few exceptions, have a complete circle of hooklets, and the larvae usually feed in some concealed situation. The pupa is incompletely obtect, with three (in some females only two) to five free abdominal segments, and emerges partly from the cocoon before the moth appears. The remaster serves to anchor the pupa to its cocoon at the correct degree of emergence, and thus facilitates the eclosion of the imago.

The Cossidae are a small family of large moths (figs. 8, 18, 19) belonging to this section, characterized by their heads with erect rough scales or hairs, the pectin ate feelers of the males, their reduced maxillae so that no food is taken in the perfect state, and their

t . -""

rs it .Q "; ' if

fill

Tim

FIG. 18.-Stygia

a u s lr a lis. S.

Europe. FIG. 19.-Zeuzera scalaris. India. wings with the fifth radial nervure arising from the third, and the main median nervure forking in the discoidal areolet. The larvae feed in plant stems, often in the wood of trees, forming tunnels and galleries, and usually taking a year or more to reach maturity. The pupa which has three or four free segments in the male and four or five in the female, rests in a cocoon within the food plant, often strengthened by chips of wood, or in a subterranean cocoon. The family is fairly well represented in the tropics; the British fauna possesses only three s cies, of which the “ goat " (Cossus cossus) and the “leopard ” f;3euzem pyrina) are well known, the caterpillars of both being often injurious to timber and fruit trees. The Tortricidae are a large family of small moths (see fig, 1), nearly allied to the Cossidae. The fifth radial nervure does not arise from the third, the maxillae are well developed, but their palps are obsolete; the head is densely clothed with erect scales; the terminal segment of the labial palp is short and obtuse. The female pupa has three, the male four, free segments. All the larvae of these moths have some method of concealing themselves while feeding. A frequent plan is to roll up a leaf of the food-plant, fastening the twisted portion with silken threads so as to make a tubular retreat; this is the habit of the caterpillar of the green bell moth (Tortrix viridana) which often ravages the foliage of oak plantations. The larvae of the pine-shoot moths (Retiwia) shelter in solidified resinous exudation's from their coniferous food-plants, while the codlin-moth caterpillar (Carpocapsa pomonella) feeds in apples and pears, growing with the growth of the fruit which affords them both provender and home. The antics of “ jumping-beans " are due to the movements of tortricid caterpillars within the substance of the seed.-The

Psychidae are a small but widely-distributed family of moths whose males have the head, densely clothed with rough hairs, bearing complex, bipectinated feelers, but with the maxillae reduced and useless. The larvae live in portable cases made of grass, pieces of leaf or stick, with a silken lining, and these cases serve as cocoons for the pupae which agree in structure with those of the Tortricidae. But the most remarkable feature of the family is the extreme degradation of the female, which, Wingless, legless and without jaws or feelers, never emerges from the cocoon. The Castniidae are a small family of large, conspicuous, day-flying exotic moths (fig.

?0) whose clubbed ' }

eelers and bright A, f

colours give them =I

a resemblance

to butterflies, al- '.

though their wing- " f*», -siéf' 1? neuration is of the

primitivetineoid A

type; the smooth larvae

feed on the - ' , ~ Mml uf'

"' 'ff rv 1 ' Nw ' ~,

stems or roots of », f4/ag, 1 lkdlliy ', plants and the I /W ~ "~m, ,

pupal structure

agrees with that of

the Tortricidae and

Psychidae. The distribution of the family is confined to Tropical America and the Indo-Malayan and Australian regions. The Zygaeuidae (burnet moths) are a large family of day-fiying moths (fig. 21) adorned with brilliant metallic colours. The feelers are long, stout in the middle and tapering, bearing numerous long or short pectinations. The well-developed maxillae have vestigial palps. The larvae-often very conspicuously coloured-are remarkable among the Tineides in having incomplete circles of hooks on the prolegs, and they feed exposed on the leaves of various plants. The f pupa, enclosed in a silken cocoon, has four or "" five free segments. The Limacodidae are a small FHL 2I Neu, 0 FIG. 20.-Castnia acraeofides. Brazil.

n 1' g, "o

i- '?>A, 'i?°

-» .. ga?

~

»-

family of brownish nocturnal moths, allied to the Zygaenidae and agreeing with them in the symploca concmnm structure of the pupa. The larva in this family S' Africa also is an exposed feeder, but it is remarkable in form, 'being fattened and slug-like, without prolegs and adorned with 'curious spinous processes.

The Sesiidae are a large family of small, narrow-winged moths, the sub-costal nervure of the hindwing being absent and the wings being for the most part

destitute of scales (fig., ¢

22). The maxillae are MV,

developed but their palps

are vestigial, while the, g " 'f-terminal segment of the ' 2,

labial palp is short and '11, , ;g », ~', pointed. Many of these ~. if

insects have their bodies . ' 5) I f 1 .ii ff banded with black and fi; f

yellow; this in conjunc- M .,

tion with the transparent f wi g *"" wings makes some of ', Q li

them like wasps or 314 » 'ii "" hornets in appearance. 35 “ The larvae feed in the T, '~.

woody stems of various A B

}§ , §§§ ;S~0rT}'§ u§ “§ ; Wx FIG. 22.-¢A, sem asimofmis (Gad-fly dominal Segments re Hawk Moth). Europe. B, Larva. mains within its cocoon, formed with chips of wood, until the time for its, final change draws near; then it works itself partly out of the tree by means of the spines on its abdominal se ments. The Nepticulidae are the smallest of all the Lepigoptera, measurmg only 3-8 mm. across the outspread wings, which are all lanceolate and pointed at the tip. The sucking portions of the maxillae are

vestigial, but the palps are long and jointed. The larvae, without